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Data:Sorex character

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SpeciesSorex albibarbis
Summer_dorsum_colorblackish or gray frosted with white
Summer_venter_colorwhite with a tinge of brownish or grayish
Winter_dorsum_color
Condylobasal_length19.1–21.7 (Beneski & Stinson, 1987); 18.4–19.2 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisors
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall
Tine_position
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shape
Dental_formula
Dental_characters
Other_characters
Tail_bicoloreddistinctly
Paired_toe_pads


SpeciesSorex arcticus
Summer_dorsum_colordark chocolate brown to blackish
Summer_venter_colorpale gray brown to whittish
Winter_dorsum_color
Condylobasal_length19.1–20.0 (Kirkland & Schmidt, 1996)
Shape_medial_edge_incisorsStraight
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall
Tine_positionWell below upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3≥U4
Postmandibular_canalPresent
Skull_shapeSkull relatively flat; braincase slightly rounded
Dental_formula3/1, 1/1, 3/1, 3/3, total = 32
Dental_charactersUnicuspids robust and appear bulbous in ventral view; tips of upper incisors are more widely separated partly because of tines
Other_characters
Tail_bicoloredfaintly
Paired_toe_pads


SpeciesSorex arizonae
Summer_dorsum_colorgrayish-brown
Summer_venter_color
Winter_dorsum_color
Condylobasal_length15.7–16.53 (Diersing & Hoffmeister, 1977); 15.7–16.53 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsStraight
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeMedium
Tine_positionWell below upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3≥U4
Postmandibular_canalPresent
Skull_shapeBraincase is slightly rounded (based on image from Diersing & Hoffmeister, 1977)
Dental_formula
Dental_charactersTips of upper incisors are more widely separated partly because of tines (Diersing & Hoffmeister, 1977)
Other_characters
Tail_bicolored
Paired_toe_pads


SpeciesSorex bendirii
Summer_dorsum_colordark brown to blackish
Summer_venter_colorslightly lighter to same color as dorsum
Winter_dorsum_color
Condylobasal_length>19.3 (Pattie, 1973); 20.8–23.8 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsSlightly curved
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeMedium
Tine_position
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shapeBraincase only slightly rounded; rostrum shallow and distinctly downcurved
Dental_formula
Dental_characters
Other_charactersSmall fringe of stiff hairs on hindfoot.
Tail_bicoloredno
Paired_toe_pads


SpeciesSorex cinereus
Summer_dorsum_colorbrown to dark grayish-brown
Summer_venter_colorgray to buffy-gray
Winter_dorsum_color
Condylobasal_length14.3–16.1 (French, 1980); 14.6–16.9 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsParallel or slightly diverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeMedium
Tine_positionWell below upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3≥U4; U1–U4 are in an almost graduated series
Postmandibular_canalAbsent
Skull_shapeRounded braincase
Dental_formula3/1, 1/1, 3/1, 3/3, total = 32
Dental_charactersTips of upper incisors are more widely separated partly because of tines
Other_characters
Tail_bicoloredfaintly
Paired_toe_pads6 on 3rd and 4th digits of HF


SpeciesSorex dispar
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length15.8–18.5 (Hall & Kelson, 1959); 16.45–18.7 (Kirkland, 1981); 15.35–18.7 (S. d. gaspensis; Kirkland, 1981); 18.4–19.2 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeTiny
Tine_positionAt or just below the upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3≥U4
Postmandibular_canalAbsent
Skull_shapeRostrum shallow; braincase bulbous; posterior border of the infraorbital foramen behind the space between the M1 and M2; mental foramen level with the space between p4 and m1
Dental_formula
Dental_characters
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex eximius
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length13.0–15.8 (Hall & Kelson, 1959); 15.0–16.2 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsSlightly curved
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeLarge and long
Tine_positionWell below upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3 and U5 are tiny and vestigial
Postmandibular_canalAbsent
Skull_shapeSkull relatively flat; braincase slightly rounded
Dental_formula
Dental_charactersU3 and U5 are barely or not at all visible in side view, but may be seen in occlusal view
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex fontinalis
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length14.63–15.85 (Kirkland & Levengood, 1987); 14.6–15.2 (Junge & Hoffman, 1981)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall
Tine_positionJust below the upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3≥U4
Postmandibular_canalAbsent
Skull_shapeSkull relatively flat
Dental_formula
Dental_characters
Other_characters
Tail_bicolored
Paired_toe_pads


SpeciesSorex fumeus
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length17.4–18.5 (Owen, 1984)
Shape_medial_edge_incisorsStrongly curved and diverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall
Tine_positionJust below the upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3>U4
Postmandibular_canalAbsent; occasionally present on one side
Skull_shapeBraincase flattened, sometimes extremely; posterior border of the infraorbital foramen ahead of the space between the M1 and M2; mental foramen approximately level with the protoconid of m1
Dental_formulai 3/1, c 1/1, p 3/1, m 3/3, total = 32
Dental_charactersWidth of unicuspids exceeds length
Other_characters
Tail_bicoloreddistinctly
Paired_toe_pads


SpeciesSorex haydeni
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length14.2–15.1 (n=10; van Zyll de Jong, 1976); 15.0–16.1 (French, 1980); 14.1–15.9 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisors
Position_upper_incisors
Tines_upper_incisorsYes
Tine_size
Tine_position
Number_unicuspids5
Unicuspid_notesU3≥U4
Postmandibular_canalAbsent
Skull_shape
Dental_formula
Dental_characters
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex hoyi
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length13.0–15.8 (Hall & Kelson, 1959); 14.3–15.0 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsSlightly curved
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeLarge and long
Tine_positionWell below upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3 and U5 are tiny and vestigial
Postmandibular_canalAbsent
Skull_shapeBraincase is rounded
Dental_formula3/1, 1/1, 3/1, 3/3, total = 32
Dental_charactersU3 and U5 are barely or not at all visible in side view, but may be seen in occlusal view
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex jacksoni
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length15.6–16.3 (Hall & Kelson, 1959)
Shape_medial_edge_incisors
Position_upper_incisors
Tines_upper_incisorsYes
Tine_size
Tine_position
Number_unicuspids5
Unicuspid_notesU3≥U4
Postmandibular_canalAbsent
Skull_shape
Dental_formula
Dental_characters
Other_characters
Tail_bicolored
Paired_toe_pads


SpeciesSorex longirostris
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length13.8–15.6, except S. l. fisheri 15.4–16.4 (French, 1980)
Shape_medial_edge_incisorsStrongly diverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall
Tine_positionAt or just below pigment line
Number_unicuspids5
Unicuspid_notesU3<U4; U3 and U4 sometimes subequal
Postmandibular_canalAbsent
Skull_shapeBraincase is rounded
Dental_formulai 3/1, c 1/1, p 3/1, m 3/3, total = 32
Dental_characters
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex lyelli
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length15.2–15.4 (Hall & Kelson, 1959)
Shape_medial_edge_incisors
Position_upper_incisors
Tines_upper_incisorsYes
Tine_size
Tine_position
Number_unicuspids5
Unicuspid_notesU3≥U4
Postmandibular_canalAbsent
Skull_shape
Dental_formula
Dental_characters
Other_characters
Tail_bicolored
Paired_toe_pads


SpeciesSorex merriami
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length15.7–17.1 (Armstrong & Jones, 1971), 16–18 (Armstrong, 2010); 15.1–16.9 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsConverging
Position_upper_incisors
Tines_upper_incisorsNo
Tine_size
Tine_position
Number_unicuspids5
Unicuspid_notesU3>U4; U2 is largest
Postmandibular_canalPresent
Skull_shapeBraincase relatively flat
Dental_formula
Dental_charactersTips of upper incisors are close together since teeth converge distally
Other_characters
Tail_bicoloreddistinctly
Paired_toe_pads


SpeciesSorex minutissimus
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length12.0–14.3 (Yudin, 1971)
Shape_medial_edge_incisors
Position_upper_incisors
Tines_upper_incisors
Tine_size
Tine_position
Number_unicuspids5
Unicuspid_notesU3>U4
Postmandibular_canalPresent
Skull_shape
Dental_formula
Dental_characters
Other_characters
Tail_bicolored
Paired_toe_pads


SpeciesSorex monticolus
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length16.1–17.7 (Hennings & Hoffman, 1977); 16.1–19.2 (Junge & Hoffman, 1981) ; 16.2–16.7 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall-medium
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shapeBraincase is rounded
Dental_formula
Dental_characters
Other_characters
Tail_bicoloredsomewhat so
Paired_toe_pads


SpeciesSorex nanus
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length13.8–14.8 (Hoffman & Owen, 1980)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeMedium
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shapeBraincase narrow and appears flattened when viewed from the side
Dental_formula
Dental_charactersA diagnostic dental measurement is the distance from the posterior border of the alveolus of the fifth unicuspid to the anterior margin of the alveolus of the incisor ranges between 1.8 and 2.4 mm.
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex navigator
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length19.1–21.7 (Beneski & Stinson, 1987); up to 22 (Armstrong, 2010)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes; although lacking in some individuals
Tine_sizeTiny
Tine_positionWell below upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3<U4; U2>U1
Postmandibular_canalAbsent
Skull_shape
Dental_formula
Dental_characters
Other_charactersFringe of stiff whitish hairs on the sides of its hind feet.
Tail_bicoloreddistinctly
Paired_toe_padsfive paired pads on digits 3 and 4


SpeciesSorex obscurus
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length16.1–17.7 (Hennings & Hoffman, 1977); 16.1–19.2 (Junge & Hoffman, 1981)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall-medium
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shapeSkull somewhat slightly flattened
Dental_formula
Dental_characters
Other_characters
Tail_bicoloredsomewhat so
Paired_toe_pads


SpeciesSorex ornatus
Summer_dorsum_colordrab brownish
Summer_venter_colorpaler than dorsum; grayish to buffy brown
Winter_dorsum_colorsimilar to summer but slightly darker sorsally and lighter ventrally
Condylobasal_length14.7–17.1 (Owen & Hoffman, 1983); 15.6–17.1 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeMedium
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shapeRelatively flat and broad and depressed interorbitally
Dental_formula
Dental_charactersUpper toothrow not crowded, with distinct triangular space between postero-medial edge of the fifth unicuspid and antero-medial edge of premolar; tips of upper incisors are more widely separated partly because of tines
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex pacificus
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length19.0–22.8 (Junge & Hoffman, 1981); 21.4–23.0 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsSlightly diverging (based on illustration in Carraway, 1985)
Position_upper_incisors
Tines_upper_incisorsNo; but may have a median ridge
Tine_size
Tine_position
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shape
Dental_formula
Dental_charactersIn ventral view, the unicuspids are wider than long
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex palustris
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length19.1–21.7 (Beneski & Stinson, 1987)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shapeRostrum straight or downcurved only slightly; braincase broad and flattened
Dental_formula
Dental_characters
Other_charactersFringe of stiff whitish hairs on the sides of its hind feet.
Tail_bicoloreddistinctly
Paired_toe_padsfive paired pads on digits 3 and 4


SpeciesSorex preblei
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length13.8–14.6 (Hoffman et al., 1969); less than or equal to 14.8 (Carraway, 1998); 13.8–15.1 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeLong~~acutely pointed
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3>U4; U1–U4 are in an almost graduated series
Postmandibular_canalAbsent
Skull_shapeSkull relatively flat
Dental_formula
Dental_characters
Other_characters
Tail_bicoloreddistinctly
Paired_toe_pads


SpeciesSorex pribilofensis
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length15.4–16.0 (Hall & Kelson, 1959)
Shape_medial_edge_incisors
Position_upper_incisors
Tines_upper_incisors
Tine_size
Tine_position
Number_unicuspids5
Unicuspid_notesU3>U4
Postmandibular_canalAbsent
Skull_shape
Dental_formula
Dental_characters
Other_characters
Tail_bicolored
Paired_toe_pads


SpeciesSorex rohweri
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length14.7–16.6 (Woodman & Fisher 2016)
Shape_medial_edge_incisorsDiverging; slightly curved
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeTiny
Tine_positionAt or above the upper pigment edge
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalPresent
Skull_shape
Dental_formula
Dental_charactersTines are are weak and often difficult to discern; pigmentation is pale or lacking (Woodman & Fisher, 2016)
Other_characters
Tail_bicoloreddistinctly
Paired_toe_padsfive paired toe pads on the 3rd 4th digits of the HF


SpeciesSorex sonomae
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_lengthS. s. sonomae: 19.34–23.22 (Carraway, 1989); S. s. tenelliodus: 16.5–20.21 (Carraway, 1989)
Shape_medial_edge_incisorsParallel
Position_upper_incisors
Tines_upper_incisorsNo
Tine_size
Tine_position
Number_unicuspids5
Unicuspid_notesU3<U4; U5 usually rectangular
Postmandibular_canalAbsent
Skull_shapeBraincase is rounded
Dental_formula
Dental_charactersUnicuspids appear somewhat bulbous; fifth unicuspid is usually rectangular and is separated from the fourth premolar by a gap (Verts & Carrawy, 1998)
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads


SpeciesSorex tenellus
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length14.5–15.3 (Hoffman & Owen, 1980)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeTiny
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent
Skull_shapeBraincase is slightly rounded (based on images)
Dental_formula
Dental_characters
Other_characters
Tail_bicolored
Paired_toe_pads


SpeciesSorex trowbridgii
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length16.8–18.8 (see George, 1989 for citations); 16.8–17.5 (Álvarez-Castañeda, 2024)
Shape_medial_edge_incisorsDiverging and curved
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall-medium
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalPresent
Skull_shapeBraincase is rounded
Dental_formula
Dental_characters
Other_characters
Tail_bicoloreddistinctly
Paired_toe_pads


SpeciesSorex tundrensis
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length17.0–18.5 (Junge & Hoffman, 1981)
Shape_medial_edge_incisorsParallel or slightly diverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeMedium
Tine_positionWell below upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3>U4
Postmandibular_canalPresent
Skull_shapeBraincase is slightly rounded
Dental_formula
Dental_characters
Other_characters
Tail_bicoloredindistinctly?
Paired_toe_pads


SpeciesSorex ugyunak
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length13.7–15.2 (Northwest Territories; van Zyll de Jong, 1976); 14.78 (n=26; van Zyll de Jong, 1976); 15.1–16.0 (northern Alaska; Bee & Hall, 1956; use with caution)
Shape_medial_edge_incisorsSlightly diverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeMedium
Tine_positionBelow upper limit of pigment on upper incisors
Number_unicuspids5
Unicuspid_notesU3≥U4
Postmandibular_canalAbsent
Skull_shapeBraincase is rounded
Dental_formula
Dental_charactersTips of upper incisors are more widely separated partly because of tines
Other_characters
Tail_bicoloreddistinctly
Paired_toe_pads


SpeciesSorex vagrans
Summer_dorsum_color
Summer_venter_color
Winter_dorsum_color
Condylobasal_length15.5–17.5 (Gillihan & Foresman, 2004)
Shape_medial_edge_incisorsDiverging
Position_upper_incisors
Tines_upper_incisorsYes
Tine_sizeSmall
Tine_positionAt or above the upper edge of pigment; commonly there is a white or pale-colored area between the tine and body of the tooth
Number_unicuspids5
Unicuspid_notesU3<U4
Postmandibular_canalAbsent; occasionally present on one side
Skull_shapeBraincase is rounded
Dental_formula
Dental_characters
Other_characters
Tail_bicoloredindistinctly
Paired_toe_pads4 or less paired toe pads on the 3rd 4th digits of the HF

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